Cylinders act as a guide for the pistons that translate the heat energy of combustion into the mechanical energy necessary to move a vehicle. Piston rings seal the piston to the cylinder to contain combustion gases and also regulate the oil distribution between the piston and cylinder wall. A cylinder head closes in the top of the cylinder forming the combustion chamber which is sealed by a head gasket (head). The head provides space for air and fuel intake valves, exhaust valves, and mounts for spark plugs and fuel injectors.

The four-stroke piston cycle of internal combustion engines starts with the piston at top of the cylinder head (top dead center or TDC) during the intake stroke. The piston moves downward in the cylinder creating a vacuum that pulls an air-fuel mix into the combustion chamber through the now open intake valve.

During the power stroke, just before the piston reaches top dead center, the spark plug fires and ignites the compressed air-fuel mixture. The resulting expansion due to combustion pushes the piston back down the cylinder toward bottom dead center.

Inline cylinder arrangements number cylinders sequentially (1, 2, 3, ...) front to rear.

The transfer case splits engine power between the front and rear axles of four-wheel drive vehicles.