Cylinders act as a guide for the pistons that translate the heat energy of combustion into the mechanical energy necessary to move a vehicle. Piston rings seal the piston to the cylinder to contain combustion gases and also regulate the oil distribution between the piston and cylinder wall. A cylinder head closes in the top of the cylinder forming the combustion chamber which is sealed by a head gasket (head). The head provides space for air and fuel intake valves, exhaust valves, and mounts for spark plugs and fuel injectors.

The electric fuel pump feeds pressurized fuel through a fuel filter to the fuel injectors via the fuel rail manifold. The fuel rail contains the fuel pressure regulator which ensures that the fuel injectors receive fuel at a consistent and known rate. Excess fuel bled off by the pressure regulator returns to the fuel tank through the fuel return line.

The cast iron exhaust manifolds collect engine exhaust gas from multiple cylinder exhaust valves and deliver it to the exhaust pipe. Exhaust manifolds can be generic or specially tuned (header pipes) to the engine. Header pipes deliver higher performance but are more expensive and less durable.

During the exhaust stroke, just before the piston reaches bottom dead center the exhaust valve opens. The resulting gases from combustion are then pushed out through the exhaust valve as the piston travels up the cylinder to top dead center, completing stroke four of the four-stroke piston cycle.

The stroke cycle of an engine is governed by the crankshaft which serves to regulate the firing order of the cylinders. All cylinders are not on the same stroke at the same time and correct firing order is important to balance engine operation and minimize vibrations. A common firing order for four-cylinder engines is 1-3-4-2 which indicates that cylinders 1 and 3 fire (power stroke)together and cylinders 4 and 2 fire together.